![]() ![]() A developmental constraint can be defined as a bias on phenotypic variability, which can be caused by “the structure, character, composition, or dynamics of the developmental system” ( 5). An alternative view holds that nonadaptive factors, such as developmental constraints, may account for the empty regions of morphospace ( 2 – 4). An extreme position suggests that natural selection is the key player responsible for the distribution of observed phenotypes. ![]() This bias in the distribution of phenotypes is explained by the interplay between different mechanisms including selection, drift, and developmental constraints ( 2 – 4). ![]() These findings highlight the potential role of development in modifying selectable variation in the evolution of Drosophila sex comb length.ĭespite the immense diversity of plausible biological forms, only a restricted range of morphospace is occupied in nature ( 1 – 4). Our results show ways in which morphologies that would be favored by natural selection are apparently impossible to achieve developmentally. As a result, when increasing sex comb length, rather than displaying a similar straight vertical shape observed in other Drosophila species, long sex combs bend because rotation is blocked by a neighboring row of bristles. We find that artificial selection changes the number of bristles per comb without a proportional change in the space available for rotation. melanogaster sex comb length on the process of rotation. Here we analyze the effect of changing D. In many Drosophila species including Drosophila melanogaster, the sex comb rotates 90° to a vertical position during development. To address this issue, we studied the sex comb, a group of modified bristles used in courtship that shows marked morphological diversity among Drosophila species. Despite extensive work on phenotypic bias, the underlying developmental mechanisms explaining why particular regions of morphological space remain unoccupied are poorly understood. Several mechanisms have been proposed to explain this bias in the distribution of phenotypes including selection, drift, and developmental constraints. "Act One was fear, and I hope Act Two will be love, but now, we are in a uncertain intermezzo.In spite of the diversity of possible biological forms observed in nature, a limited range of morphospace is frequently occupied for a given trait. "He was in the intermezzo between knowledge and belief - he'd heard the news, but it hadn't really sunk in". "Life is the intermezzo between birth and death eternal reality is before and after." So, in any of these senses, this works very well if your intent is to imply that the important parts are off the edges, and this is just a temporary thing. I had to look up entr'acte - that's "a dance, piece of music, or interlude performed between two acts of a play", or just the interval itself. From Merriam-Webster:Ģ a: a movement coming between the major sections of an extended musical work (as an opera) b : a short independent instrumental compositionģ : a usually brief interlude or diversion But, there's also intermezzo - this is a word from the arts, and has a nice ring as a metaphor. Mezzanine is prettier but does make me think "mall food court". Interspace is a word, but I agree that it's not very pretty. Therefore, I have a suggestion you almost have yourself. ![]() However, in your example of how you want to use the word, you do use "of", so maybe that's okay. There are many possibilities for space, and even specifically space between things, but I can't think of any nouns that work nicely without a preposition as well - either "between" or "of" or something. ![]()
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